Craterostigma loitense

2017年发表的新种。发表时的描述如下：

Subrosulate dwarf annual herb, leaves clustered between very short internodes, sometimes with a longer trailing internode 11 mm long between leaf clusters. Stems 4-angular, puberulous, hairs blunt-tipped. Leaves subsessile or with flattened petiole to 3 mm long; blade somewhat fleshy, adaxial surface green, abaxial surface often purple, at least towards apex, ovate to narrowly so or elliptic, 6.5–11 × 2.5–4.7 mm, base attenuate, margin subentire or minutely serrate, apex narrowed into an acute or blunt tip; midrib prominent, lateral veins inconspicuous; young leaves minutely puberulous, later glabrescent. Inflorescences axillary with flowers usually held well above the leaf rosettes; flowers solitary, ebracteolate; pedicels 4–12 mm long or rarely shorter in flower, extending to 11–22 mm long in fruit, indumentum as stems. Calyx green with purple lobes or purple extending to near the base, 3.0–4.5 mm long in flower, extending to 4–5 mm long in fruit, shortly and unevenly 5-lobed, longest (adaxial) lobe 1.0–2.2 mm long, shortest (abaxial) pair of lobes 0.4–1.25 mm long, sparsely puberulous mainly along midrib of each lobe and towards lobe apices. Corolla white with purple streaking on the limb, bilabiate; tube held erect, cylindrical, 3.0–3.3 mm long, c. 1.5 mm in diameter; lips 3.0–3.4 mm long when flattened; upper lip hooded, triangular-attenuate from broad base, 3.0–3.4 mm wide at base, apex shortly bilobed; lower lip with three rounded lobes 1.5–1.7 mm long with irregular margin, palate minutely puberulous centrally and extending into throat. Stamens two, adaxial, attached at base of upper lip, filaments 0.7–1.1 mm long, convergent, the divergent anther thecae forming a cross-shape, thecae 0.6–0.7 mm long, lower theca of each stamen with a basal tuft of long hairs; staminodes two, abaxial, attached on palate of lower lip, geniculate with a swollen yellow-puberulous boss and a recurved filament together up to 1.5 mm long when flattened, apex with a vestigial anther up to 0.25 mm long. Pistil glabrous; ovary oblong, ± 2 mm long; style slender, ± 2 mm long; stigma of two broad membranous lobes, ± 0.75 mm in diameter; disc annular but with one side incomplete. Capsule elongate, (5.0–) 8.5–12.0 × 1.5–2.5 mm, glabrous; seeds ± 0.5 × 0.4 mm, bothrospermous, with longitudinal ribs and finer transverse ribs.

Distribution:—Southern Kenya (Flora of Tropical East Africa region K6), only known from the type locality.

Etymology:—Named after the type locality of the Loita Hills.

Habitat and Ecology:—Shallow soils on quartzitic rocks at 2300 m elevation, growing under the edges of tall plants. The species is reported as poikilohydric (Agnew 2013).

Conservation status:—Craterostigma loitense is only known from the type collection. The area where it occurs is grazed and trampled, which may help to maintain the open habitat the plants require, although the substrate is so nutrient-poor that this may also limit dense woody growth (A.D.Q. Agnew, pers. comm.). As the specimens of C. loitense are so tiny, they could easily be overlooked and may well occur at other sites in the Loita Hills and beyond. In the light of the limited distribution, population and threat data on this species, it is provisionally assessed as DD (Data Deficient), following the Categories and Criteria of IUCN (2012)

A-D 生境，A、D为花期，B、C为果期。E、F：解剖的花冠；G：解剖的萼片，示雌蕊；H、I：远轴端雄蕊，具退化雄蕊；K：近轴端可育雄蕊细节。绘制：E. Fischer，A源自实物材料的照片，B-K源自主模式标本。

描述的译文：

近莲座状的矮小一年生草本，叶片聚生于非常短的节间，有时有稍长可达 11 毫米的、位于簇生叶片之间的匍匐节间。茎四棱形，被微柔毛，毛发钝头。叶近无柄或具扁平的可达 3 毫米的叶柄；叶片多少肉质，正面表面绿色，背面表面常紫色，至少尖端如此，卵形至下乱想或椭圆形，6.5-11 × 2.5-4.7 毫米，基部渐狭，边缘近全缘或稍具齿，尖端渐狭成一尖头或钝头；中脉突出，侧脉不明显；幼叶稍具柔毛，后无毛。花序野生，花常被莲座状叶片围抱；花单生，具小苞片；花梗花期长 4-12 毫米或罕见更短，果期伸长至 11-22 毫米，花梗被毛。萼片绿色具紫色裂片，或紫色部分延伸至近基部。花期长 3.0-4.5 毫米，果期延长至 4-5 毫米，具短而不平均的 5 裂，最长的（近轴端的）裂片长 1.0-2.2 毫米，最短的（远轴端的）一对裂片长 0.4-1.25 毫米，少数的毛被主要分布在每个裂片的中脉，朝向裂片尖端。花冠白色具紫色斑点，二唇形；花冠管直立，圆柱形，长 3.0-3.3 毫米，直径约 1.5 毫米；唇瓣展平时长约 3.0-3.4 毫米；上唇盔状，从基部起呈三角状楔形，基部宽 3.0-3.4 毫米，顶端短二裂；下唇具三枚圆形裂片，长 1.5-1.7 毫米，具不规则的边缘，腭部稍具毛，向中部延伸至喉部。雄蕊 2，位于近轴端，着生于上唇基部，花丝长 0.7-1.1 毫米，汇聚，分叉的药室交错呈十字形，药室长约 0.6-0.7 毫米，每枚雄蕊下方的药室基部具一束长毛；不育雄蕊 2，位于远轴端，着生于下唇的腭部，具膝状的膨大，黄色而被微柔毛，再弯曲向上延伸 1.5 毫米扁平化，尖端具花药痕迹，长可达 0.25 毫米，无毛；种子约 0.5×0.4 毫米，具小孔，有纵肋和清晰的横肋。

其他的部分就不翻译了。